Mussel Species
Genera: Alasmidonta | Anodonta | Corbicula | Dreissena | Elliptio | Lampsilis | Lasmigona | Leptodea | Ligumia | Margaritifera | Pyganodon | Strophitus | Utterbackia
Genus Alasmidonta
Family Unionidae
Of the five species in this genus known to occur in New York state, the metro area has three: A. heterodon, A. undulata and A. varicosa. The other two species have ranges that reach only western and central New York state. Of the three species in this area, A. heterodon is listed as both a state and federal endangered species in New York, New Jersey and Connecticut. It is currently known from only a few sites in the metro area. A. varicosa has declined from its formerly widespread status; it is listed as threatened in New York, and as endangered in New Jersey and Connecticut. A. undulata is regarded as a species of special concern by some workers, although it is not yet legally protected in any of the three states.
Alasmidonta heterodon (Lea, 1830)
Dwarf wedgemussel
size: 45mm (up to 55mm)
beak: two concentric ridges surrounded by two to three trapezoidal ridges on the posterior slope, beak low, rounded, and elevated slightly above the hinge line
color patterns: numerous fine color rays; periostracum yellowish-olive, green or blackish; nacre bluish white
shape: distinctive shell shape (rhomboidal to subtriangular), anterior margin curved sharply
shell features: prominent posterior ridge, small size
teeth: reverse lateral teeth (2 in the right valve, 1 in the left- unique among NY taxa)
abundance: Rare
status: US, NY, NJ, CT: endangered; IUCN: endangered
conservation challenges: sites should be sought in the lower Delaware watershed in New Jersey. If found, living specimens of this species should never be collected; in addition, dead shells of all federally endangered species cannot be collected without a permit.
N.A. distribution: New Brunswick (Canada) to North Carolina present metro distribution: NY: a short reach of the lower Neversink River (upper Delaware River watershed); NJ: Flatbrook, Pequest River, Paulins Kill, upper Delaware River; CT: not present in metro area
other regional localities: CT: present in a few Connecticut River tributaries
historical localities: NJ: Hackensack and Passaic Rivers; CT: Connecticut River watershed, Housatonic River watershed, South central coast watershed
habitat: A. heterodon is found in streams, creeks and rivers, and utilizes several hosts; it favors moderate to fast flowing water on substrates of cobble, fine gravel, or fine silt and sand, and has also been found to utilize the root systems of trees at the water's edge
hosts: Cottus bairdi (Girard, 1850) mottled sculpin; Cottus cognatus (Richardson, 1836) slimy sculpin; Etheostoma nigrum (Rafinesque, 1820) johnny darter; Etheostoma olmstedi (Storer, 1842) tesselated darter; Salmo salar (Linnaeus 1758) Atlantic salmon
Alasmidonta undulata (Say, 1817)
Triangle floater
size: 80mm
beak: heavy, coarse, uneven, with concentric sculpture; umbos extend above hinge line
color patterns: bright green color rays; nacre may be white, salmon, pink or red
shape: triangular ovate, anterior rounded, ventral margin broadly rounded
shell features: smooth, shiny yellow-brown periostracum, shell thickness decreases markedly from anterior to posterior
teeth: pseudocardinals strong, one in left valve, two in right; laterals vestigial; well-developed interdental tooth in the left valve
abundance: common
status: US, NY, CT: not legally protected; NJ: threatened; declining in parts of its N.A. range and considered by some observers to be a species of special concern in the metro area
conservation challenges: this species should be closely monitored; its two sister species are already endangered or threatened
N.A. distribution: Nova Scotia (Canada) west to the St. Lawrence River drainage, and south to Floridapresent metro distribution: NY: upper Hudson River and upper Delaware River watersheds; NJ: Hackensack River; Lamington/Raritan River, Stony Brook, Ramapo River (lower Hudson River watershed), Lubbers Run/Musconetcong River, and Paulins Kill River (upper Delaware River watershed). Pequest River, Hayne's Creek; CT: Connecticut River and Thames River watersheds
other regional localities: CT: all Connecticut watersheds except southwest coast and Hudson River watersheds
historical localities: CT: Housatonic River watershed
habitat: large creeks and small rivers, sometimes lakes; found in both slow and fast-moving water; substrate may vary from silt/sand in slow-moving water to gravel/sand in fast water
hosts: Campostoma anomalum (Rafinesque, 1820) central stoneroller; Cottus cognatus (Richardson, 1836) slimy sculpin; Etheostama flobellare (Rafinesque, 1819) fantail darter; Hypentelium nigricans (Lesueur, 1817) northern hogsucker; Lepomis gibbosus (Linnaeus, 1758) pumpkinseed; Luxilus cornutus (Mitchill, 1817) common shiner; Micropterus salmoides (Lacepede, 1802) largemouth bass; Notropis rubellus (Agassiz, 1850) rosyface shiner; Rhinichthys atratulus (Hermann, 1804) blacknose dace; Rhinichthys cataractae (Valenciennes, 1842) longnose dace; Semotilus corporalis (Mitchill, 1817)
Alasmidonta varicosa (Lamarck, 1819)
Brook floater
size: 70mm
beak: coarse, shape variable, projects slightly above hinge line
color patterns: dark green, continuous color rays; bright orange foot; nacre bluish- white with salmon in the beak cavity
shape: sub-trapezoidal to sub-ovate; anterior shell margin abruptly curved; ventral margin slightly concave
shell features: inflated, rounded posterior ridge; fine corrugations on posterior slope
teeth: thin, lamellar pseudocardinals with smooth surfaces, one each in left and right valve; laterals vestigial to absent
abundance: rare
status: US, NY, NJ, CT: endangered
conservation challenges: Legal protection for this species in New Jersey is critical. Other sites in the lower Delaware watershed should be sought.
N.A. distribution: New Brunswick (Canada) to South Carolina
present metro distribution: NY: Neversink River (upper Delaware River watershed), Shawangunk Kill (upper Hudson River watershed); NJ: north branch of the Raritan River and Stony Brook Creek (lower Hudson River watershed), Musconetcong River (upper Delaware River watershed), Lamington River, Flatbrook, Paulins Kill; CT: not present in the metro area
other regional localities: NY: upper Susquehanna River watershed; CT: Thames and Connecticut River watersheds
historical localities: NY: Housatonic River watershed, Lower Hudson River watershed, Oswego River watershed; NJ: Passaic River (lower Hudson River watershed); CT: lower Connecticut River watershed, Housatonic River watershed
habitat: confined to creeks and small rivers, typically in fast water on a substrate of stable gravel or sandy shoals
hosts: Cottus cognatus (Richardson, 1836) slimy sculpin; Lepomis gibbosus (Linnaeus, 1758) pumpkinseed; Notemigonus crysoleucas (Mitchill, 1814) golden shiner; Noturus insignis (Richardson, 1836) margined madtom; Perca flavescens (Mitchill, 1814) yellow perch; Rhinichthys atratulus (Hermann, 1804) blacknose dace; Rhinichthys cataractae (Valenciennes, 1842) longnose dace
Genus Anodonta
Family Unionidae
One species of Anodonta occurs in the metro area and adjacent New Jersey. The native species Anodonta implicata is known in the two metro area from two populations, until recently abundant in the Hudson and Delaware Rivers of New York. These are reported by local biologists to be in decline - presumably affected by the proliferation of the zebra mussel Dreissena polymorpha. The current status of A. implicata in the Connecticut River and in New Jersey portions of the Delaware River is unknown. A. implicata is not legally protected by the US or the states of the metro area.
Anodonta implicata (Say, 1829)
Alewife floater
size: 100mm (up to 125mm)
beak: double-looped sculpture not nodulous; umbos not elevated above the hinge line
color patterns: occasionally with fine rays; periostracum yellowish-brown, greenish-brown, or reddish-brown; nacre pink or purple, sometimes white
shape: sub-ellipitical and elongate; ventral margin straight
shell features: a marked increasing shell thickness from posterior to anterior below the pallial line; rounded posterior ridge
teeth: absent
abundance: uncommon
status: reportedly in decline; US, NY, NJ, CT: not legally protected
conservation challenges: evidence for this species should be sought in Connecticut
N.A. distribution: Nova Scotia (Canada) west to the St. Lawrence River drainage, and south to Florida
present metro distribution: NY: upper Hudson River and upper Delaware River watersheds; NJ: Hackensack River; Lamington/Raritan River, Stony Brook, Ramapo River (lower Hudson River watershed), Lubbers Run/Musconetcong River, and Paulins Kill River (upper Delaware River watershed). Pequest River, Hayne's Creek; CT: Connecticut River and Thames River watersheds
other regional localities: CT: all Connecticut watersheds except southwest coast and Hudson River watersheds
historical localities: CT: Housatonic River watershed
habitat: favors slow, sometimes fast running water, and also quiet standing water; substrate: cobble; known from stony river bottom on the Neversink River in the lower Delaware River watershed (NY)
hosts: Alosa pseudoharengus (Wilson, 1811) alewife; Catostomus commersoni (Lacepede, 1803) white sucker; Gasterosteus aculeatus (Linneaus, 1758) three spine stickleback; Lepomis gibbous (Linnaeus, 1758) pumpkinseed; Morone americana (Gmelin, 1789) white perch
Genus Corbicula
Family Corbiculidae
Our only representative of the family Corbiculidae is the invasive species Corbicula fluminea (Muller, 1774), the asiatic clam. Although widely distributed in the US, C. fluminea is limited in the NY metro area to several lakes on Long Island, NY, and to Twin Lakes, CT, and in NJ is present in all major waterways outside the Pine Barrens.
Corbicula fluminea (Muller, 1774)
Asiatic clam
size: 50mm, though typically smaller
beak: situated centrally, and highly placed above the hinge line
color patterns: distinguished by three brown or purple radial color bands near the umbos; periostracum brown or blackish brown; nacre colored white and purple
shape: inflated, triangular, or rounded triangular in older specimens
shell features: posterior external hinge ligament, with many coarse concentric ribs on epidermis
teeth: long, finely serrated laterals, two in the right valve and one in the left; pseudocardinals short, strong, with blunt surfaces, three per valve
abundance: uncommon
status: reportedly in decline; US, NY, NJ, CT: not legally protected
conservation challenges: presence of this invasive species should be sought in New Jersey and Connecticut
N.A. distribution: nearly all major eastern and midwestern river systems; Connecticut west to Minnesota and Iowa, south to Georgia, Oklahoma and Louisiana
present metro distribution: confined to several lakes on Long Island, NY
regional distribution: NY: all major watersheds except the upper Delaware River watershed; NJ: all major watersheds outside the Pine Barrens; CT: Connecticut, Housatonic, Thames River and South central coast watersheds; East Twin Lakes (Housatonic River watershed)
habitat: large rivers, canals and lakes on any hard substrate; some tolerance to salinity; limited tolerance to low calcium and acidity
hosts: no host required; free-swimming veliger larvae
Genus Dreissena
Family Dreissendae
Two species of the genus Dreissena occur in North America. Both are established invasives. D. bugensis is confined to the eastern Great Lakes and the St. Lawrence River, and reaches only the northern edge of the region. It does not occur in the metro area. By contrast, the zebra mussel D. polymorpha is abundant in the metro area, and is distributed widely throughout the eastern and midwestern US, reaching from Quebec (Canada) south on the Atlantic slope to Georgia, and west to Minnesota, Oklahoma and Louisiana.
Note: Although not extralimital to the metro area, D. polymorpha is described here with its sister species for convenience; it is also included among the descriptions of the other metro area species.
Dreissena bugensis (Andrusov, 1897)
Quagga mussel
size: 40mm
beak: pointed
color patterns: dark, irregular stripes
shape: outer margin evenly curved, posterior end and ventral surface rounded
shell features: valves attached by an internal hinge ligament (byssus); ventrolateral shoulder reduced or absent
teeth: absent
abundance: uncommon
status: US, NY, NJ, CT: not legally protected
conservation challenges: D. bugensis does not appear to present the problems of explosive population growth, damage to power plants and displacement of native species caused by the zebra mussel
N.A. distribution: Lake Ontario (Canada), the eastern Great Lakes, and the St. Lawrence River
present metro distribution: absent
other regional localities: NY: Lake Ontario, Lake Erie, western St. Lawrence River; CT: does not occur
habitat: large rivers and lakes with a hard substrate; this species shows some tolerance of salinity, and a limited tolerance of acidity and low calcium content
hosts: no host required; free-swimming veliger larva
Dreissena polymorpha (Pallas, 1771)
Zebra mussel
size: 40mm
beak: pointed
color patterns: dark, irregular stripes
shape: triangular in cross section, ventral surface flat, outer margin curved evenly; "D"-shaped, angled posterior end
shell features: acute shoulder laterally across ventral surface
teeth: absent; valves attached by internal hinge ligament
abundance: abundant
status: US, NY, NJ, CT: not legally protected
N.A. distribution: Quebec (Canada) and the lower St. Lawrence River west to the Great Lakes basin and Minnesota, south to Georgia on the Atlantic slope, and in the Mississippi River basin to Louisiana and Oklahoma
current US range includes the states of: AK, AL, CT, IA, IL, IN, KY, LA, MD, MI, MN, MO, MS, NJ, NY, OH, OK, PA, TN, VT, WI, WV
present metro distribution: NY: lower Hudson River watershed; CT: East Twin Lakes
other regional localities: St. Lawrence River, Lake Ontario, Lake Erie, upper Hudson River, Erie Canal to the Mohawk River (upper Hudson River watershed), Susquehanna River watershed
habitat: large rivers and lakes on any hard substrate; displays some tolerance to salinity, limited tolerance to low calcium and acidity; high tolerance to cold
hosts: no host required; free-swimming veliger larva
Genus Elliptio
Family Unionidae
Widespread and abundant, utilizing a variety of hosts, and occuring in most watersheds in the metro area, Elliptio complanata is one of our most familiar freshwater mussels. A second common and widely adapted species of Elliptio, E. dilatata, reaches western New York State drainages but is not recorded from the metro area.
Elliptio complanata (Lightfoot, 1786)
Eastern elliptio
size: 100mm (may reach 125mm)
beak: trapezoidal sculpture, low and uninflated
color patterns: fine green color rays only in young individuals; periostracum may be yellowish, brownish or blackish; may have purple nacre
shape: sub-trapeziodal, may be compressed or inflated
shell features: angular posterior ridge, prominent posterior slope
teeth: well-developed triangular pseudocardinals, two in left valve and one in right valve, with rough surfaces; long, sharp laterals
abundance: abundant, often the most abundant unionid present
status: US, NY, NJ, CT: not legally protected
conservation challenges: Careful monitoring of this and all other freshwater mussel species should be done on a regular basis
N.A. distribution: Nova Scotia (Canada) to Georgia; northern parts of the Great Lakes basin from Lake Ontario to the Lake Superior drainage
present metro distribution: throughout the metro region
other regional localities: NY: all watershed drainages except the Allegheny River, Eastern Lake Erie and the Lake Erie watershed, and most of the Niagara River basin above the falls; NJ: all watersheds; CT: all major watersheds
habitat: All types of natural freshwater bodies
hosts: Perca flavescens (Mitchill, 1814) yellow perch, is the only verified host; other, suspected hosts include Fundulus diaphanus (Lesueur, 1817) banded killifish; Lepomis cyanellus (Rafinesque, 1819) green sunfish; Lepomis humilo (Girard, 1858) orange-spotted sunfish; Micropterus salmoides (Lacepede, 1802) largemouth bass; Pomoxis annularis (Rafinesque, 1818) white crappie
Genus Lampsilis
Family Unionidae
Seven Lampsilis species are recorded from New York State, of which two are known from the metro area: Lampsilis cariosa, and L. radiata. Both have wide distributions along the Atlantic slope where they remain generally common to abundant. In the metro area however both species are much less common, and L. cariosa is listed as a species of Special Concern in Connecticut. Lampsilis abrupta, which also occurs in New York State, is Federally listed as Endangered.
Lampsilis cariosa (Say, 1817)
Yellow lampmussel
size: 100mm, rarely to 120mm
beak: weakly double-looped, moderately swollen but not much elevated
color patterns: thin green color rays absent or confined to posterior shell only
nacre: bluish-white
shape: sub-ovate with evenly rounded margin; sexually dimorphic shape
shell features: clear, glossy, yellow periostracum
teeth: two pseudocardinals in left valve and one in right; pseudocardinals and laterals well-developed but variable in shape
abundance: rare
status: US, NY: not legally protected; NJ: threatened; CT: special concern (extirpated) IUCN: endangered
conservation challenges: More needs to be known about the current distribution of this species in metro area Connecticut and New Jersey especially
N.A. distribution: Nova Scotia (Canada) to Georgia
present metro distribution: NY: Hudson River watershed, scattered other areas; NJ: Ramapo River (lower Hudson River watershed), upper Delaware River watershed; CT: not present in metro area
other regional localities: NY: upper Hudson River watershed, Susquehanna River watershed; NJ: no information; CT: may occur in Connecticut River watershed
historical localities: NY: Lake Ontario watershed, upper Delaware River watershed; NJ: Passaic and Raritan Rivers (lower Hudson River watershed); CT: lower Connecticut River watershed; Housatonic River watershed
habitat: small to large rivers with moderate to fast flow, especially in riffles. Preferred substrate sand and gravel, or shifting sands downstream from large boulders. Species of Lampsilis favor rivers, but may be found as well in streams and lakes where they tolerate a wide range of conditions. In the metro area they are most likely to be found in the Hudson and Delaware River watersheds, and some species are often among the most abundant taxa in their community
hosts: Morone americana (Gmelin, 1789) white perch; Perca flavescens (Mitchill, 1814) yellow perch
Lampsilis radiata (Gmelin, 1791)
Eastern lampmussel
size: 100mm
beak: doubled looped sculpture with umbos extending above the hinge line
color pattern: numerous dark green color rays across the shell face; periostracum yellowish to brownish green; nacre tinged with pink or salmon
shape: sub-ovate elliptical with evenly-rounded margin, sexes exhibit slight dimorphism
shell features: slightly roughened, wrinkled, greenish-yellow periostracum
teeth: long, well-developed but not heavy; two pseudocardinals in right and left valves
abundance: common
status: U.S., NY, NJ, CT: not legally protected
conservation challenges: Not a species of conservation concern, but additional populations and sites should be sought
N.A. distribution: Nova Scotia (Canada) south to South Carolina along the Atlantic drainage; Lake Ontario westward through Lake Huron and the Lake Superior basins
present metro distribution: NY: Delaware River watershed; NJ: Passaic and Ramapo River watersheds, Budd Lake (lower Delaware River watershed), Wallkill River (upper Hudson River watershed); CT: not in metro area
other regional localities: NY: Oswego River watershed, Richelieu River watershed (= Lake Champlain watershed), St. Lawrence River watershed, upper Hudson River watershed, upper Susquehanna River watershed; NJ: no information; CT: Connecticut, Housatonic, Thames River and South central coast watersheds
historical localities: NY: Lake Ontario watershed; NJ: Newton Lake/creek (lower Delaware River watershed), Morris Canal and upper Saddle River (lower Hudson River watershed)
habitat: most creeks, rivers and lakes; found in all types of flows and a wide variety of substrates but favors coarse sand and gravel
hosts: Ambloplites rupestris (Rafinesque, 1817) rock bass; Lepomis gibbosus (Linnaeus, 1758) pumpkinseed; Lepomis cyanellus (Rafinesque, 1819) bluegill; Lepomis megalotis (Rafinesque, 1820) longear sunfish; Micropterus dolomica (Lacepede, 1802) smallmouth bass; Micropterus salmoides (Lacepede, 1802) largemouth bass; Morone americana (Gmelin, 1789) white perch; Natropis ludibundus (Girard, 1856) sand shiner; Perca flavescens (Mitchill, 1814) yellow perch; Pimephales notatus (Rafinesque, 1820) bluntnose minnow; Poxomis nigromaculatus (Lesueur, 1829) black crappie
Genus Lasmigona
Family Unionidae
Lasmigona subviridis, the green floater, ranges from the Carolinas to the Hudson River basin and the St. Lawrence in New York. In New Jersey, the species once occurred in the Delaware-Raritan Canal and the Passaic, Raritan, Delaware and Pequest rivers. In 1996, a single live individual was found in Stony Brook, Mercer County. The Stony Brook occurrence represents the only reported sighting of a live green floater in the state since the early 1900's. The green floater is listed as endangered in NJ.
Lasmigona subviridis (Conrad, 1835)
Green floater
size: adult less than 65 mm, typically 55 mm
beak: projecting only slightly above hinge line, may be double looped, cavity shallow
color patterns: numerous dark green rays of varying width; periostracum dull yellow to brownish-green; nacre dull bluish-white
shape: subovate, ovate, triangle-ovate, or trapezoid
shell features: fragile, thin shell; small, rounded posterior ridge;
teeth: moderately developed but delicate, laterals vestigial; left valve with an interdental tooth, two pseudocardinal teeth and two lateral teeth; right valve with one pseudocardinal tooth and one lateral tooth
abundance: NJ: rare
status: US: not legally protected; NY: threatened; NJ: endangered; CT: does not occur
conservation challenges: this species should be sought in its former localities and in any ecologically compatible habitat in New Jersey
As an endangered species, L. subviridis may not be collected without a permit from the New Jersey Department of Environmental Protection. For details of restrictions and permits for collecting fresh water mussels and other invertebrates in the states of the metro area, please go to the Permits page.
N.A. distribution: South Carolina and Cape Fear River Basin, North Carolina to the Hudson River Basin and westward to St. Lawrence River Basin, NY
present metro distribution: NY: Susquehanna River watershed; NJ: Stony Brook, Mercer County (lower Hudson River watershed): CT: does not occur
other regional localities: none
historical localities: NY: Lake Ontario watershed, Oswego River watershed, southwestern Lake Ontario watershed, upper Hudson River watershed; NJ: Passaic and Raritan rivers (lower Hudson River watershed, Delaware and Pequest rivers; CT: did not occur
habitat: quiet, meandering parts of hydrologically stable small rivers and smaller streams, most often in slow water or pools and eddies, substrate gravelly or sandy prefers slower currents
hosts: not known, may not require host fish; direct development has been documented
Genus Leptodea
Family Unionidae
Until the early 1990's, L. ochracea, the only member of Leptodea that occurs in the metro area, was a common to abundant species in the tidal Hudson River above the salt plume, and a characteristic resident of coastal tidal waters along the eastern seaboard. It now appears to be in serious decline, possibly due to the proliferation of Dreissena polymorpha, the zebra mussel. Today considered rare in our area by some workers, it has been assigned Threatened status in Connecticut, and was described as a species of Special Concern in (Williams, et al) 1993.
Leptodea orchracea (Say, 1817)
Tidewater mucket
size: 75mm
beak: weakly double-looped sculpture rising above the hinge line; moderately swollen with deep beak cavity
color pattern: many fine dark rays on pale greenish yellow; periostracum satiny, yellow to golden brown; nacre may be pink to salmon-colored
shape: sub-ovate; sexes are dimorphic
teeth: delicate but well-developed hinge, pseudocardinals compressed and curved, two in each valve
abundance: rare
status: US, NY: not legally protected; NJ, CT: threatened
conservation challenges: Control of zebra mussel populations is probably key to the viability of L. ochracea in the metro area and throughout NY. Any populations of L. ochracea species should be reported immediately. Although the disappearance of L. ochracea is documented in NY, there is need for more information on its status in NJ and CT.
N.A. distribution: Nova Scotia (Canada) to Georgia
present metro distribution: NY: freshwater tidal Hudson River
other regional localities: NY: Grass River (northern St. Lawrence River watershed); NJ: mid, lower and upper Delaware River watershed; CT: lower Connecticut River watershed
historical localities: NJ: Passaic River (lower Hudson River watershed); CT: Housatonic River watershed
habitat: freshwater tidal rivers, standing coastal ponds including oxbows and sloughs, quiet tidal water; substrate: silt, mud
hosts: Morone americana (Gmelin, 1789) white perch
Genus Ligumia
Family Unionidae
Two species of Ligumia occur in New York state. Only L. nasuta reaches the metro area, occuring in scattered locations in the Hudson, Delaware, and Housatonic rivers. It is more common in western New York state. Although considered by some researchers to be a species of special concern, L. nasuta is not legally protected in the metro area.
Ligumia nasuta (Say, 1817)
Eastern pondmussel
size: 100mm
beak: umbos extended only slightly above the hinge line
color patterns: black, rarely with fine color rays over the posterior half of the shell; periostracum dark brown; nacre bluish-white but may be purple
shape: elongate, and pointed posteriorly
shell features: distinct posterior ridge
teeth: one to two pseudocardinals in each valve; laterals long and straight
abundance: uncommon status: US, NY: not legally protected; NJ: threatened; CT: special concern
conservation challenges: Appears to be in decline in CT; reportedly widespread but sporadic in NY; little information for NJ; increased efforts should be made to locate and monitor populations of this species, especially in NJ and CT
N.A. distribution: Nova Scotia (Canada) to South Carolina and west to the northern Interior Basin
present metro distribution: NY: upper Delaware River watershed; NJ: Swartswood Lake (upper Delaware River watershed), Delaware River, Cohansey Creek, Mantua Creek, Pennsauken Creek tributary and Raccoon Creek in Gloucester County, Pompeston Creek in Riverton (lower Delaware River watershed), Raritan River; CT: not present in metro area
other regional localities: NY: Allegheny River watershed, eastern Lake Erie watershed, Lake Ontario watershed, southwestern Lake Ontario watershed, St. Lawrence River watershed, Webatuck Creek (Housatonic River watershed), Oswego River watershed, upper Hudson River watershed; NJ: none known; CT: All watersheds except Hudson River watershed
historical localities: NY: Lake Erie watershed; NJ: Stony Brook (lower Hudson River watershed); CT: South central coast watershed
habitat: quiet standing water of estuaries, lakes and canals, and slow streams; substrate: silt and sand (varies)f
host: not known
Genus Margaritifera
Family Margaritiferidae
Formerly common in the Hackensack River and known from the Delaware River watershed, M. margaritifera today may be extirpated from the Hackensack River and is rare everywhere else in the metro area; it is apparently extirpated from New York State within the metro area, although still present in several other NY localities. It may remain common in restricted mountain stream habitats in the Appalachians (Strayer, 1994).
Margaritifera margaritifera (Linneaus, 1758)
Eastern pearlshell
size:150mm
beak: coarse, concentric sculpture
color patterns: no color rays
shape: elongate, sub-elliptical, ventral margin often arched dorsally
shell features: periostracum dark brown to black, series of small muscle attachment scars across a broad area of interior nacre
teeth: strong pseudocardinals, laterals absent
abundance: rare
status: US, NY, NJ: not legally protected; CT: special concern IUCN: endangered
conservation challenges: Absence of key host species in the lower Hudson River estuary may impact recovery of this species
N.A. distribution: New England maritimes to Connecticut, New York and Pennsylvania
present metro distribution: NY: not known; NJ: Hackensack River (lower Hudson River watershed); CT: Housatonic River watershed, Southwest coast watershed
other regional localities: NY: Hudson River watershed, Southern Adirondacks, northeastern Lake Ontario watershed, Oswego watershed, St Lawrence River watershed; NJ: not present; CT: lower Connecticut River watershed
historical localities: NY: Richelieu River watershed, upper Delaware River watershed; CT: Thames River watershed, South central and Southeast coast watersheds
habitat: cold, nutrient-poor, soft-water streams with moderate to fast flow; substrate fine, firm sand, gravel or cobble
hosts: Hucho hucho (Linneaus, 1758) huchen; Oncorhynchus clarki (Richardson, 1836) cuthroat trout; Oncorhynchus kisutch (Walbaum, 1792) coho salmon; Onchorynchus nerka (Walbaum, 1792) sockeye salmon; Onchorynchus tshawytscha (Walbaum, 1792) chinook salmon; Onchorynchus mykiss (Walbaum, 1792) rainbow trout; Phoxinus phoxinus (Linneaus, 1758) Eurasian minnow; Rhodeus sericeus (Pallas, 1776) bitterling; Salmo salar (Linnaeus, 1758) Atlantic salmon; Salmo trutta (Linnaeus, 1758) brown trout; Salvelinus fontinalis (Mitchill, 1814) brook trout
Genus Pyganodon
Family Unionidae
Pyganodon is a genus of several very similar species whose exact relationship and distribution in New York state remain unresolved. In the metro area, only Pyganodon cataracta is believed to occur. It is widespread in the upper Hudson and Delaware River watersheds, and in other areas of New Jersey, New York state, Connecticut and the Atlantic region.
Pyganodon cataracta (Say, 1817)
Eastern floater
size: 100mm (up to 150mm)
beak: sculpture of 5-7 low, evenly raised double loops elevated above hinge line
color patterns: fine green color rays (may be absent in older individuals); periostracum usually greenish
shape: elongate, sub-elliptical to sub-ovate
shell features: smooth, thin shell; nacre bluish-white
teeth: absent
abundance: abundant
status: US, NY, NJ, CT: not legally protected
conservation challenges: Healthy populations of its hosts and its adaptation to a wide range of habitats have helped Pyganodon maintain healthy populations in the metro area. As with other species that are still common and abundant, all known sites should be regularly monitored and new ones sought- particularly in New York
N.A. distribution: Saint Lawrence River (Canada) to Georgia, and west nearly to Michigan
present metro distribution: NY: upper Delaware River watershed; NJ: Passaic River (lower Hudson River watershed); CT: Housatonic River watershed
other regional localities: NY: Lake Ontario watershed, southeastern and southwestern Lake Ontario watersheds, Oswego River watershed, Housatonic River watershed, upper Hudson River watershed, upper Susquehanna River watershed; NJ: most New Jersey watersheds; CT: all watersheds
historical localities: NY: Allegheny River watershed, St. Lawrence River watershed; CT: Southeast coast watershed
habitat: slow and standing water, including oxbows and sloughs; substrate of fine sand, mud, and even silt
hosts: Ambloplites rupestris (Rafinesque, 1817) rock bass; Catostomus commersoni (Lacepede, 1803) white sucker; Cyprinus carpio (Linneaus, 1758) common carp; Gasterosteus aculeatus (Linnaeus, 1758) threespine stickleback; Lepomis gibbosus (Linnaeus, 1758) pumpkinseed; Lepomis macrochirus (Rafinesque, 1819) bluegill; Perca flavescens (Mitchill, 1814) yellow perch
Genus Strophitus
Family Unionidae
Strophitus undulatus remains a common to abundant species in most of the metro area and throughout its range. However, in New Jersey it is designated a species of special concern. It occurs from Nova Scotia to Georgia on the Atlantic slope, and west to Saskatchewan and Texas, thriving in small, slow-moving water as well as larger rivers and lakes. S. undulatus is the only species of Strophitus occurring in the Atlantic region.
Strophitus undulatus (Say, 1817)
Creeper
size: 100mm
beak: typically sharply defined, with coarse concentric sculpture slightly depressed
color patterns: fine green color rays may be apparent on posterior slope; periostracum yellowish or greenish, dark brown; nacre bicolored, white to bluish
shape: sub-ovate to sub-elliptical with posterior end blunt
shell features: shape variable; if beak sculpture is unclear, identification may be difficult
teeth: hinge teeth absent; pseudocardinals may be represented by a slight thickening of the nacre in each valve
abundance: common to abundant
status: US, NY, CT: not legally protected; NJ: special concern
conservation challenges: all freshwater mussels should be carefully monitored, and known locations surveyed on a regular basis to detect possible habitat degradation or population declines; additional populations of Strophitus and other mussels should be actively sought
N.A. distribution: Nova Scotia (Canada) west to Saskatchewan, south to Pennsylvania and the Atlantic slope to Georgia; and from Ontario southwest through the Ohio and Mississippi River interior basin drainages to Texas
present metro distribution: NY: all watersheds; NJ: lower Hudson River watershed, upper Delaware River watershed; CT: Housatonic River watershed
other regional localities: NY: Lake Ontario watershed, upper Hudson River watershed; NJ: upper Hudson River watershed; CT: Connecticut, Housatonic and Thames River watersheds
historical localities: CT: lower Connecticut River watershed, South central coast watershed
habitat: slow water of all sizes, and lakes; substrate: silt, sand, gravel and mixes
hosts: Ambloplites rupestris (Rafinesque, 1817) rock bass; Ameiurus melas (Rafinesque, 1820) black bullhead; Ameiurus natalis (Lesueur, 1819) yellow bullhead; Campostoma anomalum (Rafinesque, 1820) stoneroller; Cottus cognatus (Richardson, 1836) slimy sculpin; Cubea inconstans (Kirtland, 1841) brook stickleback; Etheostoma caeruleum (Storer, 1845) rainbow darter; Etheostoma exile (Girard, 1859) Iowa darter; Etheostoma flabellare (Rafinesque, 1819) fantail darter; Etheostoma nigrum (Rafinesque, 1820) Johnny darter; Etheostoma olmstedi (Storer, 1842) tessellated darter; Fundulus zebrinus (Jordan and Gilbert, 1883) plains killifish; Ictaluns punctatus (Rafinesque, 1818) channel catfish; Lepomis cyanellus (Rafinesque, 1819) green sunfish; Lepomis gibbosus (Linnaeus, 1758) pumpkinseed; Lepomis macrochirus (Rafinesque, 1819) bluegill; Lota lota (Linnaeus, 1758) burbat; Luxilis cornutus (Mitchill, 1817) common shiner; Micropterus dolomica (Lacepede, 1802) smallmouth bass; Micropterus salmoides (Lacepede, 1802) largemouth bass; Nocomis micropogon (Cope, 1865) river chub; Notophthalmus vidridescens (Rafinesque, 1820) red-spotted newt; Notropis hudsonius (Clinton, 1824) spottail shiner; Notropis lundibundis (Girard, 1856) sand shiner; Oncorhynchus mykiss (Waldbaum, 1792) rainbow trout; Perca flavescens (Mitchill, 1814) yellow perch; Percina caprodes (Rafinesque, 1818) log perch; Percina maculata (Girard, 1859) blackside darter; Percina phoxocephala (Nelson, 1876) slenderhead darter; Pimephales notatus (Rafinesque, 1820) bluntnose minnow; Pimephales promelas (Rafinesque, 1820) fathead minnow; Pomoxis annularis (Rafinesque, 1818) white crappie; Pomoxis nigromaculatus (Lesueur, 1829) black crappie; Rhinichthys atratulus (Hermann, 1804) blacknose dace; Rhinichthys cartaractae (Valenciennes, 1842) longnose dace; Salvelinus fontinalis (Mitchill, 1814) brook charr; Semotilus atromaculatus (Mitchill, 1818) cheek chub; Stizostedion vitreum (Mitchill, 1818) walleye; Umbra limi (Kirtland, 1840) central mudminnow; also, no host required
Genus Utterbackia
Family Unionidae
Utterbackia imbecillis is an introduced species, widespread and successful throughout central and southern North America, and now present in the Northeast region. Metro populations are apparently confined to the lower Delaware River watershed in New Jersey. It reportedly uses a wide range of host species, but can reproduce without glochidial parasitism. U. imbecillis is not legally protected by the US or the states of the metro area.
Utterbackia imbecillis (Say, 1829)
Paper pondell
size: 100mm
beak: compressed, below line of dorsal margin when viewed laterally, double looped with weak ridges
color patterns: brownish green to brown, occasional green rays; interior white to bluish white, iridescent posteriorly
shape: moderately elongate, sub-rectangular
shell features: thin and fragile, posterior pointed, hingeline straight; posterior ridge moderately angled
teeth: teeth lacking
abundance: widespread and common
status: U.S., NY, NJ, CT: not legally protected; also known as papershell or paper floater
conservation challenges: This species should be sought in other parts of the Delaware River watershed
N.A. distribution: Maryland to Wisconsin, south to Kansas, Oklahoma and Texas, and on the Atlantic slope from New York to northern Florida and Mexico
present metro distribution: NY: not present in metro area; NJ: tributaries of the lower Delaware River; CT: does not occur
other regional localities: NY: Great Lakes region; NJ: lower Delaware River tributaries;
habitat: ponds, lakes, and muddy-bottomed pools in rivers and streams
hosts: has more identified potential hosts than any other unionid, including many tropical aquarium fish and three amphibians; known fish hosts include: Ambloplites rupestris (Rafinesque, 1817) rock bass; Aplocheilus lineatus (Cuvier and Valenciennes, 1835) panehax killifish; Barbus semifasciolatus (Gunther, 1868) golden barb; Betta splendens (Ryan, 1809) Siamese fighting fish; Brachydonia kerri (Smith, 1931) blue danio; Colisa lalia (Hamilton, 1822) flame gourami; Etheostoma lepidum (Baird and Girard, 1853) greenthroat darter; Fundulus diaphanus (Lesueur, 1817) banded killifish; Gambusia affinus (Baird and Girard, 1853) mosquitofish; Haplochromis venustus (Boulenger, 1908) marbled cichlid; Hemmigrammus erythrozonus (Durbin, 1909) glowlight tetra; Lepomis cyanellus (Rafinesque, 1819) green sunfish; Lepomis gibbosus (Linnaeus, 1758) pumpkinseed; Lepomis gulosus (Cuvier, 1829) warmouth; Lepomis macrochirus (Rafinesque, 1819) bluegill; Lepomis marginatus (Holbrook, 1855) dollar sunfish; Lepomis megalotis (Rafinesque, 1820) longear sunfish; Melanotaenia maccullochi (Ogilby, 1915) Australian rainbowfish; Melynnis argenteus (Ahl, 1923) silver dollar; Micropterus salmoides (Lacepede, 1802) largemouth bass; Moenkhausia oligolepis (Gunther, 1864) redeye tetra; Pangio myers (Harry, 1949) black loach; Perca flavescens (Mitchill, 1814) yellow perch; Poecilia reticulata (Peters, 1859) guppy; Pomoxis nigromaculatus (Lesueur, 1829) black crappie; Pseudotropheus zebra (Boulenger, 1899) zebra Malawi cichlid; Rasbora einthovenii (Bleecker, 1851) brilliant rasbora; Semotrilus atromaculatus (Mitchill, 1818) creek chub; Trichogaster trichopterus (Pallas, 1770) lavender gourami; Xphophorus helleri (Heikel, 1848) painted sword
Amphibian Hosts: Ambystoma tigrinum (Green, 1825) tiger salamander; Rana catesbeiana (Shaw, 1802) bullfrog; Rana pipens (Schreber, 1782) northern leopard frog; Xenopus laevis (Daudin, 1852) African clawed frog